What makes humans unique and how did we evolve? This course will review 8 million years of hominin evolutionary history, focusing on the origins of defining features of our species including bipedalism, tool use, language, and advanced cognition. We will examine evidence from primatology and the genetic, fossil, and archaeological records. Concepts from evolutionary biology and anthropology will be covered including adaptation, phylogenetics, life history theory, behavioral ecology, and gene-culture coevolution.